tinroИзвестия ТИНРОIzvestiya TINRO1606-99192658-5510ТИНРО10.26428/1606-9919-2026-206-119-148QMYFHGtinro-1098Research ArticleБИОЛОГИЧЕСКИЕ РЕСУРСЫBIOLOGICAL RESOURCESСима Oncorhynchus masou (Salmonidae) и кижуч Oncorhynchus kisutch (Salmonidae) в бассейне Кучелиновского водохранилища (Приморский край)Masu Oncorhynchus masou (Salmonidae) and coho Oncorhynchus kisutch (Salmonidae) salmon in the ba- sin of Kuchelinovskoye Reservoir (Primorsky Region)https://orcid.org/0009-0002-1375-586XЦыгирВ. В.TsygirV. V.

Виктор Валентинович Цыгир, главный специалист

690091; пер. Шевченко, 4; Владивосток

Victor V. Tsygir, chief specialist

690091; 4, Shevchenko Alley; Vladivostok

viktor.tsygir@tinro.vniro.ruТихоокеанский филиал ВНИРО (ТИНРО)РоссияPacific branch of VNIRO (TINRO)Russian Federation2026110420262061119148Copyright © Цыгир В.В., 20262026Цыгир В.В.Tsygir V.V.Данная работа распространяется под лицензией Creative Commons Attribution 4.0.This work is licensed under a Creative Commons Attribution 4.0 License.https://izvestiya.tinro-center.ru/jour/article/view/1098

   В р. Кучелинова выше Кучелиновского водохранилища 15 июля 1987 г. были выпущены сеголетки (0+) кижуча Oncorhynchus kisutch (19,45 тыс. экз.) и симы Oncorhynchus masou (3,0 тыс. экз.). Молодь обоих видов была доставлена с Рязановского экспериментально-производственного лососевого рыборазводного завода. Приводятся результаты исследований вышеуказанных видов в 1987–1991 и 2020–2025 гг. в бассейне Кучелиновского водохранилища.

   Yearlings (0+) of coho salmon Oncorhynchus kisutch (19450 ind.) and masu salmon Oncorhynchus masou (3000 ind.) were released into the Kuchelinova River upstream of the Kuchelinovskoye Reservoir on July 15, 1987. The juveniles of both species were transported from the Ryazanovsky Experimental Salmon Hatchery. Results of these species monitoring in the basin of reservoir in 1987–1991 and 2020–2025 are presented.

симаOncorhynchus masouкижучOncorhynchus kisutchозерная формаизображения чешуивозрастчисленностьmasu salmonOncorhynchus masoucoho salmonOncorhynchus kisutchlacustrine formscale imageage of fishfish abundanceИсследование не имело спонсорской поддержкиThe study had no sponsorshipReferencesАкулин В.Н., Гавренков Ю.И., Марковцев В.Г. О нахождении жилой формы симы Oncorhynchus masu (Brevoort) (Salmonidae) в водохранилище Артемовского гидроузла (Южное Приморье) // Вопр. ихтиол. — 1984. — Т. 24, № 5. — C. 856–859. EDN: EADXFN.Akulin, V.N., Gavrenkov, Yu.I., and Markovtsev, V.G., A record of the freshwater form of Oncorhynchus masu (Brevoort) (Salmonidae) in the reservoir of the Artemovsk hydrosystem (southern Primor’e), Vopr. Ikhtiol., 1984, vol. 24, no. 5, pp. 856–859. EDN: EADXFNБерг Л.С. Карликовые самцы у рыб // Природа. — 1926. — № 8. — С. 96–100.Berg, L.S., Karlikovyye samtsy u ryb (Dwarf males in fish), Priroda, 1926, no. 8, pp. 96–100.Волобуев В.В., Марченко С.Л. Тихоокеанские лососи континентального побережья Охотского моря (биология, популяционная структура, динамика численности, промысел) : моногр. — Магадан : СВНЦ ДВО РАН, 2011. — 303 с.Volobuev, V.V. and Marchenko, S.L., Tikhookeanskiye lososi kontinental’nogo poberezh’ya Okhotskogo morya (biologiya, populyatsionnaya struktura, dinamika chislennosti, promysel) (Pacific Salmon of the Continental Coast of the Okhotsk Sea (Biology, Population Structure, Abundance Dynamics, Fishery)), Magadan: Sev.-Vost. Nauchn. Tsentr Dal’nevost. Otd. Ross. Akad. Nauk, 2011.Волобуев В.В., Рогатных А.Ю. Эколого-морфологическая характеристика кижуча Oncorhynchus kisutch (Walbaum) (Salmonidae) материкового побережья Охотского моря // Вопр. ихтиол. — 1982. — Т. 22, № 6. — С. 974–980. EDN: YVSBBU.Volobuev, V.V. and Rogatnykh, A.Yu., Ecological and morphological characteristics of coho salmon Oncorhynchus kisutch (Walbaum) (Salmonidae) from the mainland coast of the Sea of Okhotsk, Vopr. Ikhtiol., 1982, vol. 22, no. 6, pp. 974–980. EDN: YVSBBUГриценко О.Ф. Биология симы и кижуча Северного Сахалина. — М. : ВНИРО, 1973. — 40 с.Gritsenko, O.F., Biologiya simy i kizhucha Severnogo Sakhalina (Biology of masu and coho salmon in Northern Sakhalin), Moscow: VNIRO, 1973.Гриценко О.Ф. Проходные рыбы острова Сахалин (систематика, экология, промысел) : моногр. — М. : ВНИРО, 2002. — 248 с.Gritsenko, O.F., Prokhodnye ryby ostrova Sakhalin (sistematika, ekologiya, promysel) (Diadromous Fishes of Sakhalin (Systematics, Ecology, Fisheries)), Moscow: VNIRO, 2002.Двинин П.А. Озерный кижуч Oncorhynchus kisutch (Walbaum) morpha relictus nova // ДАН СССР. — 1949. — Т. 69, № 5. — С. 695–697.Dvinin, P.A., Lake coho salmon Oncorhynchus kisutch (Walbaum) morpha relictus nova, Dokl. Akad. Nauk SSSR, 1949, vol. 69, no. 5, pp. 695–697.Зорбиди Ж.Х. Кижуч азиатских стад : моногр. — Петропавловск-Камчатский : Камчат-НИРО, 2010. — 308 с.Zorbidi, J.H., Kizhuch aziatskikh stad (Coho salmon of Asian herds), Petropavlovsk-Kamchatsky: KamchatNIRO, 2010.Иванков В.Н., Броневский А.М. Постларвальная неотения у рыб // Зоол. журн. — 1978. — Т. 57, № 1. — С. 87–93.Ivankov, V.N. and Bronevsky, A.M., Postlarval neoteny in fishes, Zool. Zh., 1978, vol. 57, no. 1, pp. 87–93.Иванков В.Н., Падецкий С.Н., Карпенко С.Н., Лукьянов П.Е. Биология проходных рыб Южного Приморья // Биология проходных рыб Дальнего Востока. — Владивосток : ДВГУ, 1984. — С. 10–36. EDN: WBQWNZ.Ivankov, V.N., Padetsky, S.N., Karpenko, S.N., and Lukyanov, P.E., Biology of anadromous fishes in southern Primorsky Krai, in Biologiya prokhodnykh ryb Dal’nego Vostoka (Biology of Anadromous Fishes in the Far East), Vladivostok: Dal’nevost. Gos. Univ., 1984, pp. 10–36. EDN: WBQWNZКашкин К.А., Золотухин С.Ф., Цыгир В.В. Катадромная миграция молоди кеты в реках Южного Приморья // Современное состояние исследований лососевидных рыб : тез. докл. 3-го Всесоюз. совещ. по лососевидным рыбам. — Тольятти : ИЭВБ АН СССР, 1988. — С. 150–151.Kashkin, K.A., Zolotukhin, S.F., and Tsygir, V.V., Catadromous migration of chum salmon in the rivers of southern Primorye, in Tezisy dokl. 3-go Vsesoyuz. soveshch. po lososevidnym rybam “Sovremennoe sostoyanie issledovanii lososevidnykh ryb” (Proc. 3<sup>rd</sup> All-Sov. Meet. Salmonids “The Current State of Salmonid Research”), Tolyatti: Inst. Ekol. Volzhskogo Basseina Аkad. Nauk SSSR, 1988, pp. 150–151.Кириллова Е.А., Кириллов П.И., Малютина А.М. и др. Жилой кижуч Oncorhynchus kisutch в азиатской части ареала. К вопросу о пресноводной компоненте в структуре вида // Вопр. ихтиол. — 2021. — Т. 61, № 5. — С. 553–575. DOI: 10.31857/S0042875221050118. EDN: XRAUCU.Kirillova, E.A., Kirillov, P.I., Kuzishchin, K.V., Pavlov, D.S., Malyutina, A.M., and Gruzdeva, M.A., Resident coho salmon Oncorhynchus kisutch in the asian part of range. Revisiting the freshwater component in the structure of the species, J. Ichthyol., 2021, vol. 61, no. 5, pp. 709–730. doi: 10.1134/S003294522105009X. EDN: XRAUCUКрасная книга Камчатского края. Том 1. Животные / отв. ред. А.М. Токранов. — Петропавловск-Камчатский : Камчатпресс, 2018. — 196 с.Red Data Book of Kamchatskiy Krai. Vol. 1. Animals, Tokranov, A.M., ed., Petropavlovsk-Kamchatsky: Kamchatpress, 2018.Крыхтин М.Л. Материалы о речном периоде жизни молоди симы // Изв ТИНРО. — 1962. — Т. 48. — С. 84–132.Krykhtin, M.L., Materials on the riverine period in the life history of juvenile masu salmon, Izv. Tikhookean. Nauchno-Issled. Inst. Rybn. Khoz. Okeanogr., 1962, vol. 48, pp. 84–132.Куренков С.И. Жилой кижуч на Камчатке // Вопр. географии Камчатки. — 1977. — № 7. — С. 52–55.Kurenkov, S.I., Residential coho salmon in Kamchatka, Vopr. geografii Kamchatki, 1977, no. 7, pp. 52–55.Куренков С.И., Горшков С.А., Толстяк Т.И. Распространение и биология пресноводного кижуча Oncorhynchus kisutch (Walb.) (Salmonidae) на Камчатке // Вопр. ихтиол. — 1982. — Т. 22, № 6. — С. 966–973.Kurenkov, S.I., Gorshkov, S.A., and Tolstyak, T.I., The range and biology of the freshwater coho salmon Oncorhynchus kisutch (Walbaum) (Salmonidae) in Kamchatka, Vopr. Ikhtiol., 1982, vol. 22, no. 6, pp. 966–973.Любаев В.Я. Экологические и биотехнические аспекты создания стад кижуча на рыбоводных заводах : автореф. дис. … канд. биол. наук. — М. : МГТА, 2002. — 22 с.Lyubaev, V.Ya., Ecological and biotechnical aspects of creating coho salmon stocks at fish hatcheries, Extended Abstract of Cand. Sci. (Biol.) Dissertation, Moscow: Moskovskaya gumanitarno-tekhnicheskaya akademiya, 2002.Марченко С.Л. Кижуч Oncorhynchus kisutch (Walbaum) материкового побережья Охотского моря // Изв. ТИНРО. — 2022. — Т. 202, вып. 3. — С. 556–579. DOI: 10.26428/1606-9919-2022-202-556-579. EDN: FRJIJM.Marchenko, S.L., Coho salmon Oncorhynchus kisutch (Walbaum) on the continental coast of the Okhotsk Sea, Izv. Tikhookean. Nauchno-Issled. Inst. Rybn. Khoz. Okeanogr., 2022, vol. 202, no. 3, pp. 556–579. doi: 10.26428/1606-9919-2022-202-556-579. EDN: FRJIJMМарченко С.Л., Волобуев В.В., Макаров Д.В. Биологическая структура кижуча Oncorhynchus kisutch (Walbaum) материкового побережья Охотского моря // Исслед. водн. биол. ресурсов Камчатки и сев.-зап. части Тихого океана. — 2013. — Вып. 29. — С. 70–83.Marchenko, S.L., Volobuev, V.V., and Makarov, D.V., Biological structure of coho Oncorhynchus kisutch (Walbaum) of the continental sea-shore of the Sea of Okhotsk, Issled. Vodn. Biol. Resur. Kamchatki Sev.-Zapadn. Chasti Tikhogo Okeana, 2013, vol. 29, pp. 70–83.Моисеев П.А. Образование жилой формы симы (Oncorhynchus masou morpha formosanus (Jordan et Oshima)) в бассейне реки Седанка // ДАН СССР. — 1957. — Т. 112, № 1. — С. 163–164.Moiseev, P.A., Formation of a residential form of masu salmon (Oncorhynchus masou morpha formosanus (Jordan et Oshima)) in the Sedanka River basin, Dokl. Akad. Nauk SSSR, 1957, vol. 112, no. 1, pp. 163–164.Правдин И.Ф. Руководство по изучению рыб (преимущественно пресноводных) : моногр. — 4-е изд., перераб. и доп. — М. : Пищепромиздат, 1966. — 375 с.Pravdin, I.F., Rukovodstvo po izucheniyu ryb (preimushchestvenno presnovodnykh) (Guide to the Study of Fish (Mainly Freshwater)), 4<sup>th</sup> ed., Moscow: Pishchevaya Promyshlennost’, 1966.Самуйлов А.Е. Рыбы бассейна рек Майхе и Батальянза // Ученые записки ДВГУ. — 1971. — Т. 15, вып. 3. — С. 130–131.Samuylov, A.Ye., Fishes of the Maikhe and Batalyanza River Basins, Uchenyye zapiski Dal’nevostochnogo gosudarstvennogo universiteta, 1971, vol. 15, no. 3, pp. 130–131.Токранов А.М. Пресноводные формы лососей Камчатского края // На перекрестке континентов : мат-лы XXXI Крашенинниковских чтений. — Петропавловск-Камчатский : Камч. краев. науч. б-ка им. С.П. Крашенинникова, 2014. — С. 342–345.Tokranov, A.M., Freshwater forms of salmon in the Kamchatka region, in Na perekrestke kontinentov : mat-ly XXXI Krasheninnikovskikh chteniy (At the crossroads of continents : materials of the XXXI Krasheninnikov Readings), Petropavlovsk-Kamchatsky: Kamch. krayev. nauch. b-ka im. S.P. Krasheninnikova, 2014, pp. 342–345.Цыгир В.В. Возраст симы Oncorhynchus masou // Вопр. ихтиол. — 1988. — Т. 28, вып. 2. — С. 248–258.Tsygir, V.V., Age-related masu salmon Oncorhynchus masou, Vopr. Ikhtiol., 1988, vol. 28, no. 2, pp. 248–258.Цыгир В.В. Жилая сима из бассейна Седанкинского водохранилища (южное Приморье) // Биология шельфовых и проходных рыб. — Владивосток : ДВО АН СССР, 1990. — С. 47–52.Tsygir, V.V., Residential masu salmon from the Sedankinskoe reservoir basin (Southern Primorye), in Biologiya shel’fovykh i prokhodnykh ryb (Biology of shelf and anadromous fish), Vladivostok: Dal’nevost. Otd., Akad. Nauk. SSSR, 1990, pp. 47–52.Цыгир В.В., Барабанщиков Е.И., Шаповалов М.Е. Новые данные о симе Oncorhynchus masou (Salmonidae) из бассейна Дачинского водохранилища (Приморский край) // Изв. ТИНРО. — 2025. — Т. 205, вып. 4. — С. 625–640. DOI: 10.26428/1606-9919-2025-205-625-640. EDN: KYRFJF.Tsygir, V.V., Barabanshchikov, E.I., and Shapovalov, M.E., New data on masu salmon Oncorhynchus masou (Salmonidae) from the Dachinskoye reservoir (Primorye Region), Izv. Tikhookean. Nauchno-Issled. Inst. Rybn. Khoz. Okeanogr., 2025, vol. 205, no. 4, pp. 625–640. doi: 10.26428/1606-9919-2025-205-625-640. EDN: KYRFJFЦыгир В.В., Зайцев С.В. Выживаемость потомства проходных и неотенических самцов симы // Современное состояние исследований лососевидных рыб : тез. докл. 3-го Всесоюз. совещ. по лососевидным рыбам. — Тольятти : ИЭВБ АН СССР, 1988. — С. 356–358.Tsygir, V.V. and Zaitsev, S.V., Survival of the offspring of anadromous and neotenic male masu salmon, in Tezisy dokl. 3-go Vsesoyuz. soveshch. po lososevidnym rybam “Sovremennoe sostoyanie issledovanii lososevidnykh ryb” (Proc. 3<sup>rd</sup> All-Sov. Meet. Salmonids “The Current State of Salmonid Research”), Tolyatti: Inst. Ekol. Volzhskogo Basseina Аkad. Nauk SSSR, 1988, pp. 356–358.Цыгир В.В., Иванков В.Н. Жилая сима Oncorhynchus masou из бассейна Артемовского водохранилища (Приморский край) // Вопр. ихтиол. — 1987. — Т. 27, № 4. — С. 576–583.Tsygir, V.V. and Ivankov, V.N., The freshwater masu Oncorhynchus masou from the Artemovsk reservior basin (the Maritime Province of the USSR Pacific coast), Vopr. Ikhtiol., 1987, vol. 27, no. 4, pp. 576−583.Шедько С.В., Шедько М.Б. Новые данные по пресноводной ихтиофауне юга Дальнего Востока России // Чтения памяти Владимира Яковлевича Леванидова. — Владивосток : Дальнаука, 2003. — Вып. 2. — С. 319–336.Shedko, S.V. and Shedko, M.B., A new data on freshwater ichthyofauna of the south of the Russian Far East, in Chteniya pamyati Vladimira Yakovlevicha Levanidova (Vladimir Yakovlevich Levanidov’s Biennial Memorial Meetings), Vladivostok: Dal’nauka, 2003, vol. 2, pp. 319–336.Beamish R.J., Weitkamp L.A., Shaul L.D., Radchenko V.I. Ocean Life History of Coho Salmon // The Ocean Ecology of Pacific Salmon and Trout. — Bethesda, Maryland : American Fisheries Society, 2018. — P. 391–533.Beamish, R.J., Weitkamp, L.A., Shaul, L.D., and Radchenko, V.I., Ocean Life History of Coho Salmon, The Ocean Ecology of Pacific Salmon and Trout., Bethesda, Maryland: American Fisheries Society, 2018, pp. 391–533.Clutter R.I., Whitesel L.E. Collection and interpretation of sockeye salmon scales : Int. Pacif. Salmon Fish. Comm. — 1956. — Vol. 9. — 159 p.Clutter, R.I. and Whitesel, L.E., Collection and interpretation of sockeye salmon scales, Int. Pacif. Salmon Fish. Comm., 1956, vol. 9.Godfrey H. Salmon of the North Pacific Ocean: Part IX. Coho, chinook and masu salmon in offshore waters. 1. Coho salmon in offshore waters // Int. North Pac. Fish. Comm. Bull. — 1965. — Vol. 16. — P. 1–39.Godfrey, H., Salmon of the North Pacific Ocean: Part IX. Coho, chinook and masu salmon in offshore waters. 1. Coho salmon in offshore waters, Int. North Pac. Fish. Comm. Bull., 1965, vol. 16, pp. 1–39.Godfrey H., Henry K.A., Machidori S. Distribution and abundance of coho salmon in offshore waters of the North Pacific Ocean : Int. North Pac. Fish. Comm. Bull. — 1975. — Vol. 3. — 80 p.Godfrey, H., Henry, K.A., and Machidori, S., Distribution and abundance of coho salmon in offshore waters of the North Pacific Ocean, Int. North Pac. Fish. Comm. Bull., 1975, vol. 3.Honda N., Kataoka T., Hoshino M., Seki Y. Studies on the reproduction of landlocked masu salmon Oncorhynchus masou (Brevoot), in Okutadami reservoir. IV. Resources amount of the salmonid fishes which ascend the Kitanomata river // Rept. Niigata Pref. Inland Water Fish. Exp. St. — 1983a. — Vol. 10. — P. 1−12 (in Japanese).Honda, N., Kataoka, T., Hoshino, M., and Seki, Y., Studies on the reproduction of landlocked masu salmon Oncorhynchus masou (Brevoot), in Okutadami reservoir. IV. Resources amount of the salmonid fishes which ascend the Kitanomata river, Rept. Niigata Pref. Inland Water Fish. Exp. St., 1983, vol. 10, pp. 1−12 (in Japanese).Honda N., Kataoka T., Hoshino M., Seki Y. Studies on the reproduction of landlocked masu salmon Oncorhynchus masou (Brevoot), in Okutadami reservoir. V. Studies on the growth and age of maturity of the land-locked masu in Okutadami reservoir // Rept. Niigata Pref. Inland Water Fish. Exp. St. — 1983b. — Vol. 10. — P. 13−20 (in Japanese).Honda, N., Kataoka, T., Hoshino, M., and Seki, Y., Studies on the reproduction of landlocked masu salmon Oncorhynchus masou (Brevoot), in Okutadami reservoir. V. Studies on the growth and age of maturity of the land-locked masu in Okutadami reservoir, Rept. Niigata Pref. Inland Water Fish. Exp. St., 1983, vol. 10, pp. 13−20 (in Japanese).Honda N., Kataoka T., Hoshino M., Seki Y. Studies on the reproduction of landlocked masu salmon, Oncorhynchus masou, in Okutadami reservoir. II. Ecology of land1ocked masu salmon during the period of upstream migration for spawning // Rept. Niigata Pref. Inland Water Fish. Exp. St. — 1981. — Vol. 9. — P. 16−23 (in Japanese).Honda, N., Kataoka, T., Hoshino, M., and Seki, Y., Studies on the reproduction of landlocked masu salmon, Oncorhynchus masou, in Okutadami reservoir. II. Ecology of land1ocked masu salmon during the period of upstream migration for spawning, Rept. Niigata Pref. Inland Water Fish. Exp. St., 1981, vol. 9, pp. 16−23 (in Japanese).Honda N., Suzuki K., Amita K. et al. Studies on reproduction of landlocked masu salmon Oncorhynchus masou (Brevoot) in Okutadami reservoir. I. Vertical distribution and feeding habits of fish // Rept. Niigata Pref. Inland Water Fish. Exp. St. — 1980. — Vol. 8. — P. 5−15 (in Japanese).Honda, N., Suzuki, K., Amita, K., Kataoka, T., and Emura, K., Studies on reproduction of landlocked masu salmon Oncorhynchus masou (Brevoot) in Okutadami reservoir. I. Vertical distribution and feeding habits of fish, Rept. Niigata Pref. Inland Water Fish. Exp. St., 1980, vol. 8, pp. 5−15 (in Japanese).Kataoka T., Honda N., Suzuki A. Studies on the reproduction of landlocked masu salmon, Oncorhynchus masou, in the Okutadami reservoir — VI. Resource amount of the salmonids // Rept. Niigata Pref. Inland Water Fish. Exp. St. — 1984. — Vol. 11. — P. 1–13 (in Japanese).Kataoka, T., Honda, N., and Suzuki, A., Studies on the reproduction of landlocked masu salmon, Oncorhynchus masou, in the Okutadami reservoir — VI. Resource amount of the salmonids, Rept. Niigata Pref. Inland Water Fish. Exp. St., 1984, vol. 11, pp. 1–13 (in Japanese).MacLellan S.E. Guide for sampling structures used in age determination of Pacific salmon. — British Columbia, Nanaimo : Pacific Biological Station, 1987. — 27 p.MacLellan, S.E., Guide for sampling structures used in age determination of Pacific salmon, British Columbia, Nanaimo: Pacific Biological Station, 1987.MacLellan S.E. Guide for sampling structures used in age determination of Pacific salmon. — British Columbia, Nanaimo : Pacific Biological Station, 2004. — 31 p.MacLellan, S.E., Guide for sampling structures used in age determination of Pacific salmon, British Columbia, Nanaimo: Pacific Biological Station, 2004.Morita K. General biology of masu salmon // The Ocean Ecology of Pacific Salmon and Trout. — Bethesda, Maryland : American Fisheries Society, 2018. — P. 703–730.Morita, K., General biology of masu salmon, in The Ocean Ecology of Pacific Salmon and Trout, Bethesda, Maryland: American Fisheries Society, 2018, pp. 703–730.Osanai M. Ecological Studies on the landlocked masu salmon, Oncorhynchus masou (Brevoort). 1. Ecological succession on the limnological conditions and feeding habit of the lake-locked form at Uryu reservoir // Sci. Rep. Hok. Fish. Hatch. — 1962. — Vol. 17. — P. 21–29 (in Japanese).Osanai, M., Ecological Studies on the landlocked masu salmon, Oncorhynchus masou (Brevoort). 1. Ecological succession on the limnological conditions and feeding habit of the lake-locked form at Uryu reservoir, Sci. Rep. Hok. Fish. Hatch., 1962, vol. 17, pp. 21–29 (in Japanese).Sandercock F.K. Life history of coho salmon (Oncorhynchus kisutch) // Pacific Salmon Life Histories / eds C. Groot and L. Margolis. — Vancouver : UBC press, 1991. — P. 395–446.Sandercock, F.K., Life history of coho salmon (Oncorhynchus kisutch), Pacific Salmon Life Histories, Groot, C. and Margolis, L., eds, Vancouver: UBC Press, 1991, pp. 395–446.Tamate T., Maekawa K. Life cycle of masu salmon (Oncorhynchus masou) in Shumarinai Lake, northern Hokkaido, Japan // Eurasian J. For. Res. — 2000. — Vol. 1. — P. 39–42.Tamate, T. and Maekawa, K., Life cycle of masu salmon (Oncorhynchus masou) in Shumarinai Lake, northern Hokkaido, Japan, Eurasian J. For. Res., 2000, vol. 1, pp. 39–42.Tsiger V.V., Skirin V.I., Krupyanko N.I. et al. Life history forms of male masu salmon (Oncorhynchus masou) in South Primor’e, Russia // Can. J. Fish. Aquat. Sci. — 1994. — Vol. 51, № 1. — P. 197–208. DOI: 10.1139/f94-022.Tsiger, V.V., Skirin, V.I., Krupyanko, N.I., Kashkin, K.A., and Semenchenko, A.Yu., Life history forms of male masu salmon (Oncorhynchus masou) in South Primor’e, Russia, Can. J. Fish. Aquat. Sci., 1994, vol. 51, № 1, pp. 197–208. doi: 10.1139/f94-022Utoh H. Study of the mechanism of differentiation between the stream resident form and the sea-ward migratory form in masu salmon, Oncorhynchus masou Brevoort. I. Growth and sexual maturity of precocious masu salmon parr // Bull. Fac. Fish. Hokkaido Univ. — 1976. — Vol. 26(4). — P. 321–326 (in Japanese).Utoh, H., Study of the mechanism of differentiation between the stream resident form and the seaward migratory form in masu salmon, Oncorhynchus masou Brevoort. I. Growth and sexual maturity of precocious masu salmon parr, Bull. Fac. Fish. Hokkaido Univ., 1976, vol. 26, no. 4, pp. 321–326 (in Japanese).Utoh H. Study of the mechanism of differentiation between the stream resident form and the sea-ward migratory form in masu salmon, Oncorhynchus masou Brevoort. II. Growth and sexual maturity of precocious masu salmon parr // Bull. Fac. Fish. Hokkaido Univ. — 1977. — Vol. 28(2). — P. 66–73 (in Japanese).Utoh, H., Study of the mechanism of differentiation between the stream resident form and the seaward migratory form in masu salmon, Oncorhynchus masou Brevoort. II. Growth and sexual maturity of precocious masu salmon parr, Bull. Fac. Fish. Hokkaido Univ., 1977, vol. 28, no. 2, pp. 66–73 (in Japanese).Yamaguchi K., Nakamura T., Maruyama T. Age composition, sex ratio, growth, and feeding habit of native and introduced hatchery-reared lake-run masu salmon, Oncorhynchus masou masou in a Japanese mountain reservoir // Suisan zoshoku. — 2000. — Vol. 48, № 4. — P. 615–622 (in Japanese).Yamaguchi, K., Nakamura, T., and Maruyama, T., Age composition, sex ratio, growth, and feeding habit of native and introduced hatchery-reared lake-run masu salmon, Oncorhynchus masou masou in a Japanese mountain reservoir, Suisan zoshoku, 2000, vol. 48, no. 4, pp. 615–622 (in Japanese).Romanov, N.S., Razrabotka biologicheskikh osnov upravlyayemogo lososevogo khozyaystva na baze zavodskikh populyatsiy ryb. Materialy po vliyaniyu samtsov simy na biologicheskiye pokazateli potomstva (Development of the Biological Foundations of Managed Salmon Farming Based on Hatchery Populations. Materials on the Effect of Male Cherry Salmon on the Biological Parameters of Offspring), Available from TINRO, 1986, Vladivostok, no. GR 01823005274, inv. no. 0287.0 039753.Romanov, N.S., Razrabotka biologicheskikh osnov upravlyayemogo lososevogo khozyaystva na baze zavodskikh populyatsiy ryb. Materialy po vliyaniyu samtsov simy na biologicheskiye pokazateli potomstva (Development of the Biological Foundations of Managed Salmon Farming Based on Hatchery Populations. Materials on the Effect of Male Cherry Salmon on the Biological Parameters of Offspring), Available from TINRO, 1986, Vladivostok, no. GR 01823005274, inv. no. 0287.0 039753.Romanov, N.S., Sovershenstvovaniye biotekhniki iskusstvennogo razvedeniya tikhookeanskikh lososey. Materialy po presnovodnomu vyrashchivaniyu i adaptatsii molodi kizhucha k morskoy vode (Improving Biotechnology for Artificial Breeding of Pacific Salmon. Materials on Freshwater Cultivation and Adaptation of Juvenile Coho Salmon to Seawater), Available from TINRO, 1987, Vladivostok, no. GR 01822005315, inv. no. 028.80 017653.Romanov, N.S., Sovershenstvovaniye biotekhniki iskusstvennogo razvedeniya tikhookeanskikh lososey. Materialy po presnovodnomu vyrashchivaniyu i adaptatsii molodi kizhucha k morskoy vode (Improving Biotechnology for Artificial Breeding of Pacific Salmon. Materials on Freshwater Cultivation and Adaptation of Juvenile Coho Salmon to Seawater), Available from TINRO, 1987, Vladivostok, no. GR 01822005315, inv. no. 028.80 017653.

The authors declare that there are no conflicts of interest present.