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<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="ru"><front><journal-meta><journal-id journal-id-type="publisher-id">tinro</journal-id><journal-title-group><journal-title xml:lang="ru">Известия ТИНРО</journal-title><trans-title-group xml:lang="en"><trans-title>Izvestiya TINRO</trans-title></trans-title-group></journal-title-group><issn pub-type="ppub">1606-9919</issn><issn pub-type="epub">2658-5510</issn><publisher><publisher-name>ТИНРО</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.26428/1606-9919-2022-202-836-849</article-id><article-id custom-type="elpub" pub-id-type="custom">tinro-783</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>БИОЛОГИЧЕСКИЕ РЕСУРСЫ</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>BIOLOGICAL RESOURCES</subject></subj-group></article-categories><title-group><article-title>О валидности видов рода Cleisthenes (Pleuronectiformes: Pleuronectidae)</article-title><trans-title-group xml:lang="en"><trans-title>On validity of species of the genus Cleisthenes (Pleuronectiformes: Pleuronectidae)</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0001-9797-7763</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Бадаев</surname><given-names>О. З.</given-names></name><name name-style="western" xml:lang="en"><surname>Badaev</surname><given-names>O. Z.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Бадаев Олег Зинурович, кандидат биологических наук, ведущий научный сотрудник</p><p>690091, г. Владивосток, пер. Шевченко, 4</p></bio><bio xml:lang="en"><p>Oleg Z. Badaev - Ph.D., leading researcher</p><p>690091, Vladivostok, Shevchenko Alley, 4</p></bio><email xlink:type="simple">badayev@yandex.ru</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Черниенко</surname><given-names>И. С.</given-names></name><name name-style="western" xml:lang="en"><surname>Chernienko</surname><given-names>I. S.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Черниенко Игорь Сергеевич, кандидат биологических наук, ведущий научный сотрудник</p><p>690091, г. Владивосток, пер. Шевченко, 4</p></bio><bio xml:lang="en"><p>Igor S. Chernienko - Ph.D., leading researcher</p><p>690091, Vladivostok, Shevchenko Alley, 4</p></bio><email xlink:type="simple">Igor.chernienko@tinro-center.ru</email><xref ref-type="aff" rid="aff-1"/></contrib></contrib-group><aff-alternatives id="aff-1"><aff xml:lang="ru"><institution>Тихоокеанский филиал ВНИРО (ТИНРО)</institution><country>Россия</country></aff><aff xml:lang="en"><institution>Pacific branch of VNIRO (TINRO)</institution><country>Russian Federation</country></aff></aff-alternatives><pub-date pub-type="collection"><year>2022</year></pub-date><pub-date pub-type="epub"><day>30</day><month>12</month><year>2022</year></pub-date><volume>202</volume><issue>4</issue><fpage>836</fpage><lpage>849</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Бадаев О.З., Черниенко И.С., 2023</copyright-statement><copyright-year>2023</copyright-year><copyright-holder xml:lang="ru">Бадаев О.З., Черниенко И.С.</copyright-holder><copyright-holder xml:lang="en">Badaev O.Z., Chernienko I.S.</copyright-holder><license xml:lang="ru" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>Данная работа распространяется под лицензией Creative Commons Attribution 4.0.</license-p></license><license xml:lang="en" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://izvestiya.tinro-center.ru/jour/article/view/783">https://izvestiya.tinro-center.ru/jour/article/view/783</self-uri><abstract><p>Рассмотрена правомерность применения видовых таксонов рода Cleisthenes. На основе литературных данных показано значительное и достоверное расхождение рыб по морфологии, ставится под сомнение сведение форм «pinetorum» и «herzensteini» в один вид. Представители исследуемого рода, обитающие в водах зал. Сендай (Япония, префектуры Мияги и Фукусима), очевидно, являются Cleisthenes pinetorum. На остальной части ареала рода Cleisthenes обитает C. herzensteini.</p></abstract><trans-abstract xml:lang="en"><p>Taxonomy status of genus Cleisthenes was revised several times from the time of its description (Jordan and Starks, 1904) to nowadays. Some species of this genus were even assigned to genus Hippoglossoides. At present, the stumbling block in taxonomy of the genus is: whether the genus is represented by two species (pointhead flounder Cleisthenes herzensteini and pinewood flounder C. pinetorum) or only one. The problem cannot be easy solved because of many contradictions in previous studies. On the base of retrospective information on taxonomy of these/this species, statistical significance for divergence of some morphological indices is identified by the Kolmogorov-Smirnov criterion of homogeneity, that prevents reduction of diversity for the forms pinetorum and herzensteini to one species. The flounders of genus Cleisthenes dwelling the waters of the Sendai Bay (Miyagi and Fukushima prefectures of Japan) are obviously C. pinetorum, whereas the rest of this genus range is inhabited by C. herzensteini.</p></trans-abstract><kwd-group xml:lang="ru"><kwd>Cleisthenes</kwd><kwd>herzensteini</kwd><kwd>pinetorum</kwd><kwd>остроголовая камбала</kwd><kwd>систематика</kwd><kwd>ареал</kwd><kwd>виды</kwd><kwd>морфология</kwd></kwd-group><kwd-group xml:lang="en"><kwd>Cleisthenes</kwd><kwd>herzensteini</kwd><kwd>pinetorum</kwd><kwd>pointhead flounder</kwd><kwd>pinewood flounder</kwd><kwd>taxonomy</kwd><kwd>range of genus</kwd><kwd>species</kwd><kwd>morphology</kwd></kwd-group><funding-group><funding-statement xml:lang="ru">Авторы искренне признательны за содействие при подготовке работы профессору Музея Хоккайдского университета д-ру Ф. Тасиро.</funding-statement><funding-statement xml:lang="en">The authors are sincerely grateful to Dr. F. Tashiro, Professor of the Museum of Hokkaido University, for his assistance in the work.</funding-statement></funding-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Борец Л.А. Донные ихтиоцены российского шельфа дальневосточных морей: состав, структура, элементы функционирования и промысловое значение : моногр. — Владивосток : ТИНРО-центр, 1997. — 217 с.</mixed-citation><mixed-citation xml:lang="en">Borets, L.A., Donnye ikhtiotseny rossiiskogo shel’fa dal’nevostochnykh morei: sostav, struktura, elementy funktsionirovaniya i promyslovoye znacheniye (Benthic Ichthyocoenes on the Russian Shelf of the Far Eastern Seas: Composition, Structure, Functioning Elements, and Commercial Significance), Vladivostok: TINRO-Tsentr, 1997.</mixed-citation></citation-alternatives></ref><ref id="cit2"><label>2</label><citation-alternatives><mixed-citation xml:lang="ru">Булатов Н.В., Самко Е.В., Цыпышева И.Л. Океанологические образования, благоприятные для концентрации пелагических рыб по инфракрасным данным ИСЗ NOAA // Современные проблемы дистанционного зондирования Земли из космоса. — 2008. — Т. 2, вып. 5. — С. 49–61.</mixed-citation><mixed-citation xml:lang="en">Bulatov, N.V., Samko, E.V., and Tsypysheva, I.L., Oceanological formations favorable for the detection of pelagic fish according to NOAA satellites, Sovremennyye problemy distantsionnogo zondirovaniya Zemli iz kosmosa, 2008, vol. 2, no. 5, pp. 49–61.</mixed-citation></citation-alternatives></ref><ref id="cit3"><label>3</label><citation-alternatives><mixed-citation xml:lang="ru">Булатов О.А., Котенев Б.Н., Кровнин А.С. О перспективах новой «сардиновой эпохи» в северо-западной части Тихого океана // Вопр. рыб-ва. — 2016. — Т. 17, № 4. — С. 385–405.</mixed-citation><mixed-citation xml:lang="en">Bulatov, O.A., Kotenev, B.N., and Krovnin, A.S., On the prospects for a new “Sardine era” in the northwestern Pacific, Vopr. Rybolov., 2016, vol. 17, no. 4, pp. 385–405.</mixed-citation></citation-alternatives></ref><ref id="cit4"><label>4</label><citation-alternatives><mixed-citation xml:lang="ru">Воронина Е.П. Особенности строения сейсмосенсорной системы некоторых представителей семейства Pleuronectidae (sensu Chapleau, Keast, 1988). Сообщение 1 // Вопр. ихтиол. — 2002. — Т. 42, № 5. — С. 581–590.</mixed-citation><mixed-citation xml:lang="en">Voronina, E.P., Morphology of the Acoustico-Lateralis System of Some Pleuronectids: Communication 1, J. Ichthyol., 2002, vol. 42, no. 8, pp. 555–564.</mixed-citation></citation-alternatives></ref><ref id="cit5"><label>5</label><citation-alternatives><mixed-citation xml:lang="ru">Воронина Е.П. Особенности строения сейсмосенсорной системы некоторых представителей семейства Pleuronectidae (sensu Chapleau, Keast, 1988). Сообщение 3 // Вопр. ихтиол. — 2005. — Т. 45, № 2. — С. 157–167.</mixed-citation><mixed-citation xml:lang="en">Voronina, E.P., Characteristics of the Seismosensory System Morphology in Some Pleuronectidae, J. Ichthyol., 2005, vol. 45, no. 2, pp. 146–156.</mixed-citation></citation-alternatives></ref><ref id="cit6"><label>6</label><citation-alternatives><mixed-citation xml:lang="ru">Долганов В.Н. Формирование биологического разнообразия дальневосточных красноперок рода Tribolodon (Cyprinidae) // Биол. моря. — 2021. — Т. 47, № 6. — С. 369–380. DOI:10.31857/S013434752106005X.</mixed-citation><mixed-citation xml:lang="en">Dolganov, V.N., The formation of biological diversity of far eastern redfins of the genus Tribolodon (Cyprinidae), Russ. J. Mar. Biol., 2021, vol. 47, no. 6, pp. 369–380. doi:10.31857/S013434752106005X</mixed-citation></citation-alternatives></ref><ref id="cit7"><label>7</label><citation-alternatives><mixed-citation xml:lang="ru">Дьяков Ю.П. Камбалообразные (Pleuronectiformes) дальневосточных морей России (пространственная организация, сезоны и продолжительность нереста, популяционная структура вида, динамика популяций) : моногр. — Петропавловск-Камчатский : КамчатНИРО, 2011. — 433 с.</mixed-citation><mixed-citation xml:lang="en">Dyakov, Yu.P., Kambaloobraznye (Pleuronectiformes) dal’nevostochnykh morei Rossii (Flatfish (Pleuronectiformes) of the Far Eastern Seas of Russia), Petropavlovsk-Kamchatsky: KamchatNIRO, 2011.</mixed-citation></citation-alternatives></ref><ref id="cit8"><label>8</label><citation-alternatives><mixed-citation xml:lang="ru">Линдберг Г.У., Федоров В.В. Рыбы Японского моря и сопредельных частей Охотского и Желтого морей : моногр. — СПб. : Наука, 1993. — Ч. 6. — 272 с. (Определители по фауне СССР, изд. АН СССР, № 166.)</mixed-citation><mixed-citation xml:lang="en">Lindberg, G.U. and Fedorov, V.V., Opredeliteli po faune, izdavaemye zoologicheskim institutom Rossiiskoi akademii nauk. T. 166. Ryby Yaponskogo morya i sopredel’nykh chastei Okhotskogo i Zheltogo morei (Keys to Fauna Published by the Zoological Institute, Russian Academy of Sciences, vol. 166: Fishes of the Sea of Japan and Adjacent Waters of the Sea of Okhotsk and the Yellow Sea), St. Petersburg: Nauka, 1993, part 6.</mixed-citation></citation-alternatives></ref><ref id="cit9"><label>9</label><citation-alternatives><mixed-citation xml:lang="ru">Парин Н.В., Евсеенко С.А., Васильева Е.Д. Рыбы морей России: аннотированный каталог. — М. : Тов-во науч. изд. КМК, 2014. — 733 с.</mixed-citation><mixed-citation xml:lang="en">Parin, N.V., Evseenko, S.A., and Vasiljeva, E.D., Ryby morei Rossii: annotirovannyi katalog (Fishes of Russian Seas: Annotated Catalogue), Moscow: KMK, 2014.</mixed-citation></citation-alternatives></ref><ref id="cit10"><label>10</label><citation-alternatives><mixed-citation xml:lang="ru">Редин А.Д., Картавцев Ю.Ф. Филогенетические отношения камбалообразных рыб семейства Pleuronectidae (Ostichties: Pleuronectiformes) на основе участка гена 16S рРНК // Генетика. — 2021. — Т. 57, № 3. — С. 345–357. DOI:10.31857/S0016675821030115.</mixed-citation><mixed-citation xml:lang="en">Redin, A.D. and Kartavtsev, Y.P., Phylogenetic relationships of flounders from the family Pleuronectidae (Ostichties: Pleuronectiformes) based on 16s rRNA gene, Rus. J. Genet., 2021, vol. 57, no. 3, pp. 345–357. doi:10.31857/S0016675821030115</mixed-citation></citation-alternatives></ref><ref id="cit11"><label>11</label><citation-alternatives><mixed-citation xml:lang="ru">Соколовский А.С., Соколовская Т.Г., Яковлев Ю.М. Рыбы залива Петра Великого : моногр. — 2-е изд., испр. и доп. — Владивосток : Дальнаука, 2011. — 431 с.</mixed-citation><mixed-citation xml:lang="en">Sokolovsky, A.S., Sokolovskaya, T.G., and Yakovlev, Yu.M., Ryby zaliva Petra Velikogo (Fishes of Peter the Great Bay), Vladivostok: Dal’nauka, 2011.</mixed-citation></citation-alternatives></ref><ref id="cit12"><label>12</label><citation-alternatives><mixed-citation xml:lang="ru">Фадеев Н.С. Северотихоокеанские камбалы (распространение и биология) : моногр. — М. : Агропромиздат, 1987. — 175 с.</mixed-citation><mixed-citation xml:lang="en">Fadeev, N.S., Severotikhookeanskie kambaly (rasprostranenie i biologiya) (North Pacific Flounders (Distribution and Biology)), Moscow: Agropromizdat, 1987.</mixed-citation></citation-alternatives></ref><ref id="cit13"><label>13</label><citation-alternatives><mixed-citation xml:lang="ru">Федосова Р.А. Межгодовые изменения мезопланктона в районе к юго-востоку от побережья Японии // Изменчивость состава ихтиофауны, урожайности поколений и методы прогнозирования запасов рыб в северной части Тихого океана. — Владивосток : ТИНРО, 1988. — С. 77−83.</mixed-citation><mixed-citation xml:lang="en">Fedosova, R.A., Interannual changes in mesoplankton in the area southeast of the coast of Japanб, in Izmenchivost’ sostava ikhtiofauny, produktivnost’ pokoleniy i metody prognoza rybnykh zapasov severnoy chasti Tikhogo okeana (Variability in the composition of the ichthyofauna, productivity of generations and methods for forecasting fish stocks in the northern part of the Pacific Ocean), Vladivostok: TINRO, pp. 77–83.</mixed-citation></citation-alternatives></ref><ref id="cit14"><label>14</label><citation-alternatives><mixed-citation xml:lang="ru">Шитиков В.К., Розенберг Г.С., Зинченко Т.Д. Количественная гидроэкология: опыт системной идентификации : моногр. — Тольятти : ИЭВБ РАН, 2003. — 463 c.</mixed-citation><mixed-citation xml:lang="en">Shitikov, V.K., Rozenberg, G.S., and Zinchenko, T.D., Kolichestvennaya gidroekologiya: opyt sistemnoy identifikatsii (Quantitative hydroecology: experience of system identification), Tolyatti: IEVB RAN, 2003.</mixed-citation></citation-alternatives></ref><ref id="cit15"><label>15</label><citation-alternatives><mixed-citation xml:lang="ru">Шмидт П.Ю. Рыбы восточных морей Российской империи : моногр. — СПб. : Изд-во Император. рус. геогра. о-ва, 1904. — 466 с.</mixed-citation><mixed-citation xml:lang="en">Shmidt, P.Yu., Ryby vostochnykh morei Rossiiskoi imperii (Fishes from the Eastern Seas of the Russian Empire), St. Petersburg: Izd. Imp. Russ. Geogr. O-va, 1904.</mixed-citation></citation-alternatives></ref><ref id="cit16"><label>16</label><citation-alternatives><mixed-citation xml:lang="ru">Шмидт П.Ю. Рыбы Охотского моря : моногр. — М. ; Л. : АН СССР, 1950. — 370 с.</mixed-citation><mixed-citation xml:lang="en">Schmidt, P.Yu., Ryby Okhotskogo morya (Fishes of the Sea of Okhotsk), Moscow: Akad. Nauk SSSR, 1950.</mixed-citation></citation-alternatives></ref><ref id="cit17"><label>17</label><citation-alternatives><mixed-citation xml:lang="ru">Cheng Q.T. Pleuronectiformes. — China : Shandong Science and Technology Press, 1997. — 549 p. (In Chinese).</mixed-citation><mixed-citation xml:lang="en">Cheng, Q.T., Pleuronectiformes, China: Shandong Science and Technology Press, 1997.</mixed-citation></citation-alternatives></ref><ref id="cit18"><label>18</label><citation-alternatives><mixed-citation xml:lang="ru">Choi Y., Kim J.H., Park J.Y. Marine fishes of Korea. — Seoul : Kyohak Publishing, 2002. — 747 p. (In Korean).</mixed-citation><mixed-citation xml:lang="en">Choi, Y., Kim, J.H., and Park, J.Y., Marine fishes of Korea, Seoul: Kyohak Publishing, 2002.</mixed-citation></citation-alternatives></ref><ref id="cit19"><label>19</label><citation-alternatives><mixed-citation xml:lang="ru">Cooper J.A., Chapleau F. Monophyly and intrarelationships of the family Pleuronectidae (Pleuronectiformes), with a revised classification // Fish. Bull. — 1998. — Vol. 96(4). — P. 686–726.</mixed-citation><mixed-citation xml:lang="en">Cooper, J.A. and Chapleau, F., Monophyly and intrarelationships of the family Pleuronectidae (Pleuronectiformes), with a revised classification, Fish. Bull., 1998, vol. 96(4), pp. 686–726.</mixed-citation></citation-alternatives></ref><ref id="cit20"><label>20</label><citation-alternatives><mixed-citation xml:lang="ru">Dyldin Y.V., Orlov A.M., Ichthyofauna of fresh and brackish waters of Sakhalin Island: An annotated list with taxonomic comments: 4. Pholidae–Tetraodontidae families // J. Ichthyol. — 2017. — Vol. 57, № 2. — P. 183–218. DOI:10.1134/S0032945217020072.</mixed-citation><mixed-citation xml:lang="en">Dyldin, Y.V. and Orlov, A.M., Ichthyofauna of fresh and brackish waters of Sakhalin Island: An annotated list with taxonomic comments: 4. Pholidae–Tetraodontidae families, J. Ichthyol., 2017, vol. 57, no. 2, pp. 183–218. doi:10.1134/S0032945217020072</mixed-citation></citation-alternatives></ref><ref id="cit21"><label>21</label><citation-alternatives><mixed-citation xml:lang="ru">Dyldin Yu.V., Fricke R., Hanel L. et al. Freshwater and brackish water fishes of Sakhalin Island (Russia) in inland and coastal waters: an annotated checklist with taxonomic comments // Zootaxa. — 2021. — Vol. 5065, № 1. — P. 1–92. DOI:10.11646/zootaxa.5065.1.1.</mixed-citation><mixed-citation xml:lang="en">Dyldin, Yu.V., Fricke, R., Hanel, L., Vorobiev, D.S., Interesova, E.A., Romanov, V.I., and Orlov A.M., Freshwater and brackish water fishes of Sakhalin Island (Russia) in inland and coastal waters: an annotated checklist with taxonomic comments, Zootaxa, 2021, vol. 5065, no. 1, pp. 1–92. doi:10.11646/zootaxa.5065.1.1</mixed-citation></citation-alternatives></ref><ref id="cit22"><label>22</label><citation-alternatives><mixed-citation xml:lang="ru">Evseenko S.A. Family Pleuronectidae Cuvier, 1816 — righteye flounders : Calif. Acad. Sci. Annotated checklists of fishes. — San Francisco, 2004. — № 37. — 37 p.</mixed-citation><mixed-citation xml:lang="en">Evseenko, S.A., Family Pleuronectidae Cuvier, 1816 — righteye flounders, Calif. Acad. Sci. Annotated checklists of fishes, San Francisco, 2004, no. 37.</mixed-citation></citation-alternatives></ref><ref id="cit23"><label>23</label><citation-alternatives><mixed-citation xml:lang="ru">Fishes of Japan: with pictorial keys and species / ed. T. Nakabo. — Tokyo : Tokai Univ. Press, 2002. — Vol. 1–2. — 1749 p.</mixed-citation><mixed-citation xml:lang="en">Fishes of Japan: with Pictorial Keys and Species, Nakabo, T., ed., Tokyo: Tokai Univ. Press, 2002, vol. 1–2.</mixed-citation></citation-alternatives></ref><ref id="cit24"><label>24</label><citation-alternatives><mixed-citation xml:lang="ru">Garrett D.L., Pietsch T.W., Utter F.M., Hauser L. The Hybrid Sole Inopsetta ischyra (Teleostei: Pleuronectiformes: Pleuronectidae): Hybrid or Biological Species? // Trans. Am. Fish. Soc. — 2007. — Vol. 136, Iss. 2. — P. 460–468. DOI:10.1577/T06-092.1.</mixed-citation><mixed-citation xml:lang="en">Garrett, D.L., Pietsch, T.W., Utter, F.M., and Hauser, L., The Hybrid Sole Inopsetta ischyra (Teleostei: Pleuronectiformes: Pleuronectidae): Hybrid or Biological Species?, Trans. Am. Fish. Soc., 2007, vol. 136, no. 2, pp. 460–468. doi:10.1577/T06-092.1</mixed-citation></citation-alternatives></ref><ref id="cit25"><label>25</label><citation-alternatives><mixed-citation xml:lang="ru">Hubbs C.L. Flounders and soles from Japan collected by the United States Bureau of Fisheries steamer «Albatross» in 1906 // Proc. U.S. Natn. Mus. — 1915. — Vol. 48, № 2082. — P. 449–496. DOI:10.5479/SI.00963801.48-2082.449.</mixed-citation><mixed-citation xml:lang="en">Hubbs, C.L., Flounders and soles from Japan collected by the United States Bureau of Fisheries steamer “Albatross” in 1906, Proc. U.S. Natn. Mus., 1915, vol. 48, no. 2082, pp. 449–496. doi:10.5479/si.00963801.48-2082.449</mixed-citation></citation-alternatives></ref><ref id="cit26"><label>26</label><citation-alternatives><mixed-citation xml:lang="ru">Inada T., Murakami M. Fluctuations of Pollock and Pacific cod resources and bottom temperature in the waters off the Tohoku region of Japan // Sci. Rep. Hokkaido Exp. Stn. — 1993. — Vol. 42. — P. 229–240. (In Japanese).</mixed-citation><mixed-citation xml:lang="en">Inada, T. and Murakami, M., Fluctuations of walleye pollock and Pacific cod resources and bottom temperature in the waters off the Tohoku region of Japan, Sci. Rep. Hokkaido Exp. Stn., 1993, vol. 42, pp. 229–240 (in Japanese).</mixed-citation></citation-alternatives></ref><ref id="cit27"><label>27</label><citation-alternatives><mixed-citation xml:lang="ru">Jordan D.S., Hubbs C.L., McGregor E.A. Record of fishes obtained in Japan by David Starr Jordan in Japan, 1922 // Mem. Carnegie Mus. — 1925. — Vol. 10, № 2. — P. 93–346. DOI:10.5962/p.234844.</mixed-citation><mixed-citation xml:lang="en">Jordan, D.S., Hubbs, C.L., and McGregor, E.A., Record of fishes obtained in Japan by David Starr Jordan in Japan, 1922, Mem. Carnegie Mus., 1925, vol. 10, no. 2, pp. 93–346. doi:10.5962/p.234844</mixed-citation></citation-alternatives></ref><ref id="cit28"><label>28</label><citation-alternatives><mixed-citation xml:lang="ru">Jordan D.S., Starks E.C. A review of the flounders and soles of Japan // Proceedings of the United States National Museum. — 1906. — Vol. 31(1484). — P. 161–246. DOI:10.5479/si.00963801.31-1484.161.</mixed-citation><mixed-citation xml:lang="en">Jordan, D.S. and Starks, E.C., A review of the flounders and soles of Japan, Proceedings of the United States National Museum, 1906, vol. 31(1484), pp. 161–246. doi:10.5479/si.00963801.31-1484.161</mixed-citation></citation-alternatives></ref><ref id="cit29"><label>29</label><citation-alternatives><mixed-citation xml:lang="ru">Jordan D.S., Starks E.C. List of fishes dredged by the steamer Albatross off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae // Bull. U.S. Fish Comm. — 1904. — Vol. 22. — P. 577–630.</mixed-citation><mixed-citation xml:lang="en">Jordan, D.S. and Starks, E.C., List of fishes dredged by the steamer Albatross off the coast of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae, Bull. U.S. Fish Comm., 1904, vol. 22, pp. 577–630.</mixed-citation></citation-alternatives></ref><ref id="cit30"><label>30</label><citation-alternatives><mixed-citation xml:lang="ru">Jordan D.S., Tanaka S., Snyder J.O. A catalogue of the fishes of Japan // J. Coll. Sci. Tokyo Imperial Univ. — 1913. — Vol. 33. — P 1–497.</mixed-citation><mixed-citation xml:lang="en">Jordan, D.S., Tanaka, S., and Snyder, J.O., A catalogue of the fishes of Japan, J. Coll. Sci. Tokyo Imperial Univ., 1913, vol. 33, pp. 1–497.</mixed-citation></citation-alternatives></ref><ref id="cit31"><label>31</label><citation-alternatives><mixed-citation xml:lang="ru">Kim I.-S., Youn C.-H. Taxonomic revision of the flounders (Pisces: Pleuronectidae) from Korea // Korean J. Ichthyol. — 1994. — Vol. 6, № 2. — P. 99–132. (In Korean).</mixed-citation><mixed-citation xml:lang="en">Kim, I.-S. and Youn, C.-H., Taxonomic revision of the flounders (Pisces: Pleuronectidae) from Korea, Korean J. Ichthyol., 1994, vol. 6, no. 2, pp. 99–132 (in Korean).</mixed-citation></citation-alternatives></ref><ref id="cit32"><label>32</label><citation-alternatives><mixed-citation xml:lang="ru">Kitada S., Yoshikai R., Fujita T. et al. Population structure and persistence of Pacific herring following the Great Tohoku earthquake // Conserv. Genet. — 2017. — Vol. 18. — P. 423–437. DOI:10.1007/s10592-016-0918-2.</mixed-citation><mixed-citation xml:lang="en">Kitada, S., Yoshikai, R., Fujita, T., Hamasaki, K., Nakamichi, R., and Kishino, H., Population structure and persistence of Pacific herring following the Great Tohoku earthquake, Conserv. Genet., 2017, vol. 18, pp. 423–437. doi:10.1007/s10592-016-0918-2</mixed-citation></citation-alternatives></ref><ref id="cit33"><label>33</label><citation-alternatives><mixed-citation xml:lang="ru">Kuronuma K. Cleisthenes pinetorum vs. Protopsetta herzensteini // Bull. Biogeogr. Soc. Jpn. — 1939. — Vol. 9, № 10. — P. 181–192.</mixed-citation><mixed-citation xml:lang="en">Kuronuma, K., Cleisthenes pinetorum vs. Protopsetta herzensteini, Bull. Biogeogr. Soc. Jpn., 1939, vol. 9, no. 10, pp. 181–192.</mixed-citation></citation-alternatives></ref><ref id="cit34"><label>34</label><citation-alternatives><mixed-citation xml:lang="ru">Li S.-Z., Wang H.-M. Fauna Sinica. Osteichthyes. Pleuronectiformes. — Beijing : Science Press, 1995. — 433 p. (In Chinese).</mixed-citation><mixed-citation xml:lang="en">Li, S.-Z. and Wang, H.-M., Fauna Sinica. Osteichthyes. Pleuronectiformes, Beijig: Science Press, 1995 (in Chinese).</mixed-citation></citation-alternatives></ref><ref id="cit35"><label>35</label><citation-alternatives><mixed-citation xml:lang="ru">Nelson J.S., Grande T.C., Wilson M.V.H. Fishes of the world. — Fifth edition. — Hoboken : John Wiley &amp; Sons, 2016. — 707 p. DOI:10.1002/9781119174844.</mixed-citation><mixed-citation xml:lang="en">Nelson, J.S.,Fishes of the world, 5&lt;sup&gt;rd&lt;/sup&gt; ed., Hoboken: John Wiley &amp; Sons, 2016. doi:10.1002/9781119174844</mixed-citation></citation-alternatives></ref><ref id="cit36"><label>36</label><citation-alternatives><mixed-citation xml:lang="ru">Norman J.R. A systematic monograph of the flatfishes (Heterosomata). — London : British Mus.(Nat. Hist.), 1934. — 459 p.</mixed-citation><mixed-citation xml:lang="en">Norman, J.R., A systematic monograph of the flatfishes (Heterosomata), London: British Mus. (Nat. Hist.), 1934.</mixed-citation></citation-alternatives></ref><ref id="cit37"><label>37</label><citation-alternatives><mixed-citation xml:lang="ru">Okada Y., Uchida K., Matsubara K. Color atlas of fishes of Japan. — Tokyo : Sanseido Co Ltd., 1935. — 426 p.</mixed-citation><mixed-citation xml:lang="en">Okada, Y., Uchida, K., and Matsubara, K., Color atlas of fishes of Japan, Tokyo: Sanseido Co Ltd., 1935.</mixed-citation></citation-alternatives></ref><ref id="cit38"><label>38</label><citation-alternatives><mixed-citation xml:lang="ru">Sakamoto K. Interrelationships of the family Pleuronectidae (Pisces: Pleuronectiformes) // Memoirs of the Faculty of Fisheries Hokkaido University. — 1984. — Vol. 31, № 1, 2. — P. 95–215.</mixed-citation><mixed-citation xml:lang="en">Sakamoto, K., Interrelationships of the family Pleuronectidae (Pisces: Pleuronectiformes), Memoirs of the Faculty of Fisheries, Hokkaido University, 1984, vol. 31, no. 1, 2, pp. 95–215.</mixed-citation></citation-alternatives></ref><ref id="cit39"><label>39</label><citation-alternatives><mixed-citation xml:lang="ru">Schmidt P.Yu. On a collection of flat–fishes from Fusan (Korea) // Doklady Akademii Nauk SSSR. — 1931. — Vol. 12. — P. 313–320.</mixed-citation><mixed-citation xml:lang="en">Schmidt, P.Yu., On a collection of flat–fishes from Fusan (Korea), Doklady Akademii Nauk SSSR, 1931, vol. 12, pp. 313–320.</mixed-citation></citation-alternatives></ref><ref id="cit40"><label>40</label><citation-alternatives><mixed-citation xml:lang="ru">Shinohara G., Nakae M., Ueda Y. et al. Annotated Checklist of Deep-sea Fishes of the Sea of Japan // Deep-sea Fauna of the Sea of Japan. — 2014. — № 44. — P. 225–291.</mixed-citation><mixed-citation xml:lang="en">Shinohara, G., Nakae, M., Ueda, Y., Kojima, S., and Matsuura, K., Annotated Checklist of Deep-sea Fishes of the Sea of Japan, Deep-sea Fauna of the Sea of Japan, 2014, no. 44, pp. 225–291.</mixed-citation></citation-alternatives></ref><ref id="cit41"><label>41</label><citation-alternatives><mixed-citation xml:lang="ru">Shinohara G., Narimatsu Y., Hattori T. et al. Annotated Checklist of Deep-Sea Fishes from the Pacific Coast off Tohoku District, Japan // Nat. Mus. Nat. Sci. Monographs. — 2009. — № 39. — P. 683–735.</mixed-citation><mixed-citation xml:lang="en">Shinohara, G., Narimatsu, Y., Hattori, T., Ito, M., Takata, Y., and Matsuura, K., Annotated Checklist of Deep-Sea Fishes from the Pacific Coast off Tohoku District, Japan, Nat. Mus. Nat. Sci. Monogr., 2009, vol. 39, pp. 683–735.</mixed-citation></citation-alternatives></ref><ref id="cit42"><label>42</label><citation-alternatives><mixed-citation xml:lang="ru">Shinohara G., Shirai A.M., Nazarkin M.V., Yabe M. Preliminary list of the deep-sea fishes of the Sea of Japan // Bull. Nat. Mus. Nat. Sci., Ser. A. — 2011. — Vol. 37(1). — P. 35–62.</mixed-citation><mixed-citation xml:lang="en">Shinohara, G., Shirai, A.M., Nazarkin, M.V., and Yabe, M., Preliminary list of the deep-sea fishes of the Sea of Japan, Bull. Nat. Mus. Nat. Sci., Ser. A., 2011, vol. 37(1), pp. 35–62.</mixed-citation></citation-alternatives></ref><ref id="cit43"><label>43</label><citation-alternatives><mixed-citation xml:lang="ru">Snyder J.O. Japanese shore fishes collected by the United States Bureau of Fisheries steamer «Albatross» expedition of 1906 // Proc. U.S. Natn. Mus. — 1912. — Vol. 42. — P. 399–450.</mixed-citation><mixed-citation xml:lang="en">Snyder, J.O., Japanese shore fishes collected by the United States Bureau of Fisheries steamer “Albatross” expedition of 1906, Proc. U.S. Natn. Mus., 1912, vol. 42, pp. 399–450.</mixed-citation></citation-alternatives></ref><ref id="cit44"><label>44</label><citation-alternatives><mixed-citation xml:lang="ru">Sonoyama T., Ogimoto K., Hori S. et al. An annotated checklist of marine fishes of the Sea of Japan off Yamaguchi Prefecture, Japan, with 74 new records // Bull. of the Kagoshima University Museum. — 2020. — Vol. 11. — P. 1–152. (In Japanese).</mixed-citation><mixed-citation xml:lang="en">Sonoyama, T., Ogimoto, K., Hori, S., Uchida, Y., and Kawano, M., An annotated checklist of marine fishes of the Sea of Japan off Yamaguchi Prefecture, Japan, with 74 new records, Bull. of the Kagoshima University Museum, 2020, vol. 11, pp. 1–152 (in Japanese).</mixed-citation></citation-alternatives></ref><ref id="cit45"><label>45</label><citation-alternatives><mixed-citation xml:lang="ru">Tanaka S. Fishes of Japan. — Tokio : Dai Nihon Tosho Kabushiki Kaisha, 1936. — Vol. 1–2. — 334 p. (In Japanese).</mixed-citation><mixed-citation xml:lang="en">Tanaka, S.,Fishes of Japan, Tokio: Dai Nihon Tosho Kabushiki Kaisha, 1936, Vol. 1–2 (in Japanese).</mixed-citation></citation-alternatives></ref><ref id="cit46"><label>46</label><citation-alternatives><mixed-citation xml:lang="ru">Tashiro F. What Is known of fish diversity in the Sea of Japan? Flatfishes: a case study // Fish diversity of Japan. Evolution, zoogeography and conservation. — Singapore : Springer, 2022. — P. 79–109.</mixed-citation><mixed-citation xml:lang="en">Tashiro, F., What Is known of fish diversity in the Sea of Japan? Flatfishes: a case study, Fish diversity of Japan. Evolution, zoogeography and conservation, Singapore: Springer, 2022, pp. 79–109.</mixed-citation></citation-alternatives></ref><ref id="cit47"><label>47</label><citation-alternatives><mixed-citation xml:lang="ru">Tohkairin A., Hamatsu T., Yoshikawa A. et al. An illustrated and annotated checklist of fishes on Kitami-Yamato Bank, southern Sea of Okhotsk // Publ. Seto Mar. Biol. Lab. — 2015. — Vol. 43. — P. 1–29. DOI:10.5134/193238.</mixed-citation><mixed-citation xml:lang="en">Tohkairin, A., Hamatsu, T., Yoshikawa, A., Kai, Y., and Narabo, T., An illustrated and annotated checklist of fishes on Kitami-Yamato Bank, southern Sea of Okhotsk, Publ. Seto Mar. Biol. Lab., 2015, vol. 43, pp. 1–29. doi:10.5134/193238</mixed-citation></citation-alternatives></ref><ref id="cit48"><label>48</label><citation-alternatives><mixed-citation xml:lang="ru">Vinnikov K.A., Thomson R.C., Munroe T.A. Revised classification of the righteye flounders (Teleostei: Pleuronectidae) based on multilocus phylogeny with complete taxon sampling // Molecular Phylogenetics and Evolution. — 2018. — Vol. 125. — P. 147–162. DOI:10.1016/j.ympev.2018.03.014.</mixed-citation><mixed-citation xml:lang="en">Vinnikov, K.A., Thomson, R.C., and Munroe, T.A., Revised classification of the righteye flounders (Teleostei: Pleuronectidae) based on multilocus phylogeny with complete taxon sampling, Molecular Phylogenetics and Evolution, 2018, vol. 125, pp. 147–162. doi:10.1016/j.ympev.2018.03.014</mixed-citation></citation-alternatives></ref><ref id="cit49"><label>49</label><citation-alternatives><mixed-citation xml:lang="ru">Wang S.A, Wang Z.M., Li G.L., Cao Y.P. The fauna of Hebei, China (Pisces). — Shijiazhuang : Hebei Science and Technology Publishing House, 2001. — 366 p. (In Chinese).</mixed-citation><mixed-citation xml:lang="en">Wang, S.A., Wang, Z.M., Li, G.L., and Cao, Y.P., The fauna of Hebei, China (Pisces), Shijiazhuang: Hebei Science and Technology Publishing House, 2001.</mixed-citation></citation-alternatives></ref><ref id="cit50"><label>50</label><citation-alternatives><mixed-citation xml:lang="ru">Wu H.-W. Contribution a l’étude morphologique, biologique et systématique des poissons hétérosomes (Pisces Heterosomata) de la Chine // Theses Univ. Paris. — 1932. — Vol. 244. — P. 1–179.</mixed-citation><mixed-citation xml:lang="en">Wu, H.W., Contribution a l’étude morphologique, biologique et systématique des poissons hétérosomes (Pisces Heterosomata) de la Chine, Theses Univ. Paris, 1932, vol. 244, pp. 1–179.</mixed-citation></citation-alternatives></ref><ref id="cit51"><label>51</label><citation-alternatives><mixed-citation xml:lang="ru">Fricke, R., Eschmeyer, W.N., and Van der Laan, R., Eschmeyer’s catalog of fishes: genera, species, references, 2022. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp. Electronic version accessed 27 June 2022.</mixed-citation><mixed-citation xml:lang="en">Fricke, R., Eschmeyer, W.N., and Van der Laan, R., Eschmeyer’s catalog of fishes: genera, species, references, 2022. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp. Electronic version accessed 27 June 2022.</mixed-citation></citation-alternatives></ref><ref id="cit52"><label>52</label><citation-alternatives><mixed-citation xml:lang="ru">Kaschner, K., Kesner-Reyes, K., Garilao, C., Rius-Barile, J., Rees, T. and Froese, R., AquaMaps: predicted range maps for aquatic species, 2016, World wide web electronic publication, www.aquamaps.org, Version 08/2016. Cited August 27, 2022.</mixed-citation><mixed-citation xml:lang="en">Kaschner, K., Kesner-Reyes, K., Garilao, C., Rius-Barile, J., Rees, T. and Froese, R., AquaMaps: predicted range maps for aquatic species, 2016, World wide web electronic publication, www.aquamaps.org, Version 08/2016. Cited August 27, 2022.</mixed-citation></citation-alternatives></ref></ref-list><fn-group><fn fn-type="conflict"><p>The authors declare that there are no conflicts of interest present.</p></fn></fn-group></back></article>
